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There are a number of active members on the YouTube Developer Relations team here including Jeff Posnick, Jarek Wilkiewicz, and Ibrahim Ulukaya who all have knowledge of Youtube internals... UPDATE: Increased the quota numbers to reflect current limits on December 10, 2013.
Both weekly upload quota and total storage usage are determined by the size of the original source file at the time of upload (for example, a 200MB source file will use 200MB of the weekly upload quota or total storage limit).